G. it. Ent., 10 (51): 275-300
October 2003
Studies on the Coprophagous Scarab Beetles from East Asia. VIII
Revision of the subgenus Macronthophagus of Onthophagus
(Coleoptera, Scarabaeidae)
by
TERUO OCHI
Abstract - Macronthophagus (a subgenus of Onthophagus) is revised. Two new species and one new subspecies of O. (M.) uenoi Ochi are described: O. (M.) menieri sp. nov. from Java, O. (M.) cludtsi sp. nov. from Vietnam and South China, O. (M.) uenoi borneomontanus subsp. nov. from Borneo. In addition, O. (M.) nilgirensis Gillet, previously synonymized with O. (M.) diabolicus Harold, is regarded as a full species. A key to the species and subspecies of Macronthophagus is also presented.
